Out of the 6 million active Twitter users, the researchers picked the top 100 users in each of the three categories [Number of followers, number of retweets, and number of mentions]. Due to the overlap, there were only 233 distinct users on these lists. These were dubbed the “all-time influentials.” Some of these accounts belonged to news organizations or celebs, but others were just regular users. Regarding that last group - it appears that those users who limit their tweets to a single topic are the most likely to increase their influence scores.
In the end, what the researchers found was that follower count alone is not necessarily a worthy measure of determining influence. Other factors come into play as well. Although some heavily-followed accounts are also mentioned and retweeted a lot, just looking at audience size doesn’t reveal an account’s ability to influence and impact the Twitter universe.